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Monday, April 3rd, 2017
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7:47a |
[Invertebrate • 2017] Pygmarrhopalites dbari & P. kovali • Two New Troglobiont Pygmarrhopalites Species of the principalis group (Collembola: Arrhopalitidae) from the West Caucasus
 | Pygmarrhopalites kovali
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Abstract
Springtails of the principalis-group of the genus Pygmarrhopalites from the W Caucasian caves in Abkhazia are described: Pygmarrhopalites dbari sp. nov. from Psyrtskha Cave in Novy Afon and Pygmarrhopalites kovali sp. nov. from caves of Tsebelda Karst Massif. They differ from epigean relatives mainly by troglomorphies: reduced pigmentation, elongated appendages and modified foot complex. These new species as well as a great portion of endemic Caucasian speleofauna have highly restricted distribution and require protection. A new record of Arrhopalites abchasicus Vargovitsh, 2013 in Novoafonskaya Cave is added.
Keywords: Collembola, springtails, Symphypleona, taxonomy, new record, troglomorphic, Caucasus Mountains, Abkhazia
 | Pygmarrhopalites kovali, habitus of mounted male. |
Robert S. Vargovitsh. 2017. Two New Troglobiont Pygmarrhopalites Species of the principalis group (Collembola: Arrhopalitidae) from the West Caucasus. Zootaxa. 4250(1); 23-42. DOI: 10.11646/zootaxa.4250.1.2
| 10:12a |
[Entomology • 2017] Digitonthophagus Balthasar, 1959: Taxonomy, Systematics, and Morphological Phylogeny of the Genus Revealing An African Species Complex (Coleoptera: Scarabaeidae: Scarabaeinae)  | FIGURES 7–10. Digitonthophagus spp., Male dorsal habitus. 7. Digitonthophagus bonasus, alloreferent (Tamil Nadu State, India); 8. D. uks, holotype; 9. D. sahelicus, holotype; 10. D. catta, alloreferent (Tamil Nadu State, India).
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Abstract
The taxonomy and systematics of the genus Digitonthophagus Balthasar (Coleoptera: Scarabaeidae: Scarabaeinae: Onthophagini) is revised. A detailed study of the male genitalia combined with external morphology suggests that the variability, previously recognized, for D. gazella is hiding a species complex within the Afrotropical region and the Arabian Peninsula. The current study recognizes 16 species; 13 from the Afrotropical region and Arabian Peninsula and three from the eastern portion of the Saharo-Arabian region and the continental Indomalayan region. Species are organized into six species groups based on the results of the morphology-based phylogenetic analysis. The following 12 species are described as new: D. aksumensis Génier new species; D. biflagellatus Génier new species; D. dilatatus Génier new species; D. eucatta Génier new species; D. falciger Génier new species; D. fimator Génier new species; D. namaquensis Génier new species; D. petilus Génier new species; D. sahelicus Moretto new species; D. uks Génier new species; D. ulcerosus Génier new species; and D. viridicollis Génier new species. In order to stabilize nomenclature, lectotypes are designated for Scarabaeus bonasus Fabricius, 1775; Scarabaeus catta Fabricius, 1787, and Onthophagus gazella lusinganus d’Orbigny. A neotype is designated for Scarabaeus dorcas Olivier, 1789 whose status and synonymy need to be altered in order to clarify the status of Scarabaeus gazella auctorum, the widely introduced species with economic importance. A naming scheme is presented for the sclerites of the internal sac. External and male genitalia are illustrated and distribution maps are provided for each species.
Keywords: Coleoptera, Scarabaeidae, Scarabaeinae, Digitonthophagus, Onthophagus, Digitonthophagus gazella, Nomenclature, taxonomy, distribution, Afrotropical region, Indomalayan region, biological control of dung, pasture improvement
Philippe Moretto and François Génier. 2017. Digitonthophagus Balthasar, 1959: Taxonomy, Systematics, and Morphological Phylogeny of the Genus Revealing An African Species Complex (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa. 4248(1); 1-110. DOI: 10.11646/zootaxa.4248.1.1
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