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Thursday, August 24th, 2017
Time |
Event |
2:15a |
[Entomology • 2017] Nososticta nicobarica • A New Species of Damselfly (Odonata: Platycnemididae: Disparoneurinae) from Great Nicobar Island, India
 | Nososticta nicobarica
Rajeshkumar, Raghunathan & Chandra, 2017
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Abstract
The damselfly Nososticta nicobarica sp. nov. (Odonata: Platycnemididae: Disparoneurinae) is described (holotype ♂: ZSI-ANRC-T-4418; 11.ii.2016) from Gandhi Nagar, Great Nicobar Island in the Andaman & Nicobar Islands, India. This is the first record for the genus Nososticta from the Indian sub-continent, and a formal description is given. The new species is described on the basis of male anal appendages, particularly cerci with a deeply bifurcate, acute apex, distinctly curved downward. Paraprocts almost straight with blunt apex slightly curved upward and inner margin with small, roughly rectangular hook. The female is characterised by the anterior lobe of the pronotum, which bears two long, divergent horn-like processes and a short posterior lobe with paired club-shaped processes. Affinities with species of Lesser Sundaic region are discussed.
Keywords: Odonata, Nososticta nicobarica sp. nov., Disparoneurinae, Great Nicobar Island, India
S. Rajeshkumar, C. Raghunathan and Kailash Chandra. 2017. Nososticta nicobarica sp. nov. (Odonata: Platycnemididae: Disparoneurinae) from Great Nicobar Island, India. Zootaxa. 4311(3); 426–434. DOI: 10.11646/zootaxa.4311.3.8
| 2:50a |
[Entomology • 2017] Society Islands Beach Bum Black Flies, Simulium (Inseliellum) (Diptera: Simuliidae)
 | Simulium (Inseliellum) littopyga, Simulium (Inseliellum) littosocius, Simulium (Inseliellum) littosodalis
Craig & Evenhuis, 2017 |
Abstract
Aspects of the body of work on the Central and Western Pacific black fly Simulium (Inseliellum) are briefly reviewed. Female adults collected from beaches in Tahiti and Raiatea are described as Simulium (Inseliellum) littopyga n. sp., Simulium (Inseliellum) littosocius n. sp. and Simulium (Inseliellum) littosodalis n. sp.. Immature stages of the three species are not associated.
Keywords: Diptera, Simuliidae, Simulium, Inseliellum, Society Islands, new species
Simulium (Inseliellum) littopyga Craig & Evenhuis, n. sp.
Etymology. In reference to inhabiting beaches; deriving from litto [= “beach”] + pyga [= “rump”]; hence a “beach bum” of sorts. The name is treated as a noun in apposition. Distribution. Known only from Tahiti.
Simulium (Inseliellum) littosocius Craig & Evenhuis, n. sp.
Etymology. In reference to the association with beaches; deriving from litto [= "beach"] + socius [= “companion”]; hence a second “beach bum” of sorts. The name is treated as a noun in apposition. Distribution. Known from Tahiti and Raiatea.
Simulium (Inseliellum) littosodalis Craig & Evenhuis, n. sp.
Etymology. In reference to occurring with other beach simuliids; deriving from litto [= “beach”] + sodalis [= “comrade”, “crony”]; hence a third “beach bum” of sorts. The name is treated as a noun in apposition. Distribution. Known only from Tahiti.
 | FIGURE 42. Type locality of Simulium littosodalis. Tahiti Nui, North Road, PK 42.7. 17 July 2006.
Image N. Evenhuis. |
Douglas A. Craig and Neal L. Evenhuis. 2017. Society Islands Beach Bum Black Flies (Diptera: Simuliidae). Zootaxa. 4311(3); 373–388. DOI: 10.11646/zootaxa.4311.3.3
| 4:23a |
[PaleoEntomology • 2017] Mesembrinella caenozoica • First Fossil of An Oestroid Fly (Diptera: Calyptratae: Oestroidea) and the Dating of Oestroid Divergences
 | Mesembrinella caenozoica
Cerretti, Stireman, Pape, O'Hara, Marinho, Rognes & Grimaldi, 2017
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Abstract
Calyptrate flies include about 22,000 extant species currently classified into Hippoboscoidea (tsetse, louse, and bat flies), the muscoid grade (house flies and relatives) and the Oestroidea (blow flies, bot flies, flesh flies, and relatives). Calyptrates are abundant in nearly all terrestrial ecosystems, often playing key roles as decomposers, parasites, parasitoids, vectors of pathogens, and pollinators. For oestroids, the most diverse group within calyptrates, definitive fossils have been lacking. The first unambiguous fossil of Oestroidea is described based on a specimen discovered in amber from the Dominican Republic. The specimen was identified through digital dissection by CT scans, which provided morphological data for a cladistic analysis of its phylogenetic position among extant oestroids. The few known calyptrate fossils were used as calibration points for a molecular phylogeny (16S, 28S, CAD) to estimate the timing of major diversification events among the Oestroidea. Results indicate that: (a) the fossil belongs to the family Mesembrinellidae, and it is identified and described as Mesembrinella caenozoica sp. nov.; (b) the mesembrinellids form a sister clade to the Australian endemic Ulurumyia macalpinei (Ulurumyiidae) (McAlpine’s fly), which in turn is sister to all remaining oestroids; (c) the most recent common ancestor of extant Calyptratae lived just before the K–Pg boundary (ca. 70 mya); and (d) the radiation of oestroids began in the Eocene (ca. 50 mya), with the origin of the family Mesembrinellidae dated at ca. 40 mya. These results provide new insight into the timing and rate of oestroid diversification and highlight the rapid radiation of some of the most diverse and ecologically important families of flies.

 | Fig 1. Holotype of Mesembrinella caenozoica sp. nov. (A) habitus in right dorsolateral view. (B) head and part of thorax in right dorsolateral view. (C) thorax in right dorsolateral view. |
Systematics
Order Diptera Superfamily Oestroidea
Family Mesembrinellidae Genus Mesembrinella Giglio-Tos, 1893.
Mesembrinella caenozoica sp. nov.
Type material. Holotype male, a Dominican amber inclusion of Miocene age, housed in the American Museum of Natural History. Additional details are given under Materials and Methods.
Etymology. The specific epithet ‘caenozoica’ alludes to the name of the Cenozoic Era (from Greek kainos, meaning ‘new’, and zoe, meaning ‘life’), which covers the period from ca. 66 mya to the present day. The epithet should be treated as a Latin adjective.
Pierfilippo Cerretti , John O. Stireman III, Thomas Pape, James E. O’Hara, Marco A. T. Marinho, Knut Rognes and David A. Grimaldi. 2017. First Fossil of An Oestroid Fly (Diptera: Calyptratae: Oestroidea) and the Dating of Oestroid Divergences. PLoS ONE. 12(8); e0182101. DOI: 10.1371/journal.pone.0182101
| 3:25p |
[PaleoMammalogy • 2017] Anatoliadelphys maasae • Skeleton of An Unusual, Cat-sized Marsupial Relative (Metatheria: Marsupialiformes) from the middle Eocene (Lutetian) of Turkey  | Anatoliadelphys maasae Maga & Beck, 2017
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Abstract
We describe a near-complete, three-dimensionally preserved skeleton of a metatherian (relative of modern marsupials) from the middle Eocene (Lutetian: 44–43 million years ago) Lülük member of the Uzunçarşıdere Formation, central Turkey. With an estimated body mass of 3–4 kg, about the size of a domestic cat (Felis catus) or spotted quoll (Dasyurus maculatus), it is an order of magnitude larger than the largest fossil metatherians previously known from the Cenozoic of the northern hemisphere. This new taxon is characterised by large, broad third premolars that probably represent adaptations for hard object feeding (durophagy), and its craniodental morphology suggests the capacity to generate high bite forces. Qualitative and quantitative functional analyses of its postcranial skeleton indicate that it was probably scansorial and relatively agile, perhaps broadly similar in locomotor mode to the spotted quoll, but with a greater capacity for climbing and grasping. Bayesian phylogenetic analysis of a total evidence dataset comprising 259 morphological characters and 9kb of DNA sequence data from five nuclear protein-coding genes, using both undated and “tip-and-node dating” approaches, place the new taxon outside the marsupial crown-clade, but within the clade Marsupialiformes. It demonstrates that at least one metatherian lineage evolved to occupy the small-medium, meso- or hypo-carnivore niche in the northern hemisphere during the early Cenozoic, at a time when there were numerous eutherians (placentals and their fossil relatives) filling similar niches. However, the known mammal fauna from Uzunçarşıdere Formation appears highly endemic, and geological evidence suggests that this region of Turkey was an island for at least part of the early Cenozoic, and so the new taxon may have evolved in isolation from potential eutherian competitors. Nevertheless, the new taxon reveals previously unsuspected ecomorphological disparity among northern hemisphere metatherians during the first half of the Cenozoic.
Systematic palaeontology
Mammalia; Theria Metatheria; Marsupialiformes
Anatoliadelphys gen. nov. Anatoliadelphys maasae sp. nov.
Etymology: Anatolia (Greek): the geographic name for the Asian part of Turkey; delphys (Greek): uterus, a common suffix for marsupials and their fossil relatives; maasae: in honour of Dr. Mary Maas and her contributions to Paleogene mammalian palaeontology, particularly in Turkey.
Holotype: Ankara Üniversitesi Jeoloji Müzesi (AÜJM) specimen 2002–25, which comprises a fragmented partial cranium, both dentaries, and associated postcranial elements, including most of the vertebral column, partial pectoral and pelvic girdles, all of the long limb bones, both calcanei, two metapodials, and a few phalanges.
Locality and age: AÜJM 2002–25 was collected from the Lülük member of the Uzunçarşıdere Formation (UCF), which is part of the small Orhaniye-Güvenç sedimentary basin located at the northwestern edge of the city of Ankara, approximately 5 km southwest of the town of Kazan, in central Turkey. The Lülük member is the lowest of the three members currently recognised within the UCF (together with the Gökdere [middle], and Sarıbeyler [upper] members), and is the source of all fossil mammals known from the UCF to date. AÜJM 2002–25 is from locality AK33, which is approximately 90m above the base of the UCF, at Memlik village. Until recently, the age of the UCF was poorly constrained, but a combination of U-Pb dating of zircons and magnetostratigraphy now support a date of 44–42 MYA (= Lutetian) for the formation as a whole, and 44–43 MYA for the Lülük member.
Diagnosis: Anatoliadelphys maasae differs from all other metatherians in the following combination of features: comparatively large size (estimated body mass 3–4 kg); premolars increase markedly in size posteriorly (occlusal area of p1 less than one sixth that of p3); P3 and p3 very large (similar in occlusal area to M2 and m2 respectively) and also broad (labiolingual width:mesiodistal length ratio is 0.89 for P3 and 0.7 for p3); modified tribosphenic molar dentition, in which M1-3 and m1-4 increase markedly in size posteriorly (occlusal area of M1 approximately one third that of M3; occlusal area of m1 approximately one seventh that of m4); upper molars with cingula extending along the anterior and posterior margins; protocone large but conules indistinct or absent; metacone taller than the paracone on M3 but smaller than the paracone on M4; centrocrista v-shaped on M3, with the premetacrista extending labially to stylar cusp D; centrocrista straight on M4; parastylar lobe very large on M4; anterior cingulid weakly developed on m3-4; m4 trigonid dominated by enormous protoconid, with paraconid and metaconid both greatly reduced; preentocristid and cristid obliqua of m3-4 both with carnassial notch; posterior cingulid present but very faint on m3-4; strongly curved radius and tibia; femur with prominent third trochanter, well-marked trochlea and distal condyles of approximately equal width; calcaneus with medially-inflected tuber, large peroneal process with prominent groove for peroneus longus tendon, concave calcaneocuboid facet, and prominent pit (probably for plantar calcaneocuboid ligament) on ventral surface.
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 | Fig 1. Holotype skeleton of Anatoliadelphys maasae (AÜJM 2002–25). Scale bar = 5 cm. |
 | Reconstruction of the Anatoliadelphys maasae.
Illustration: Peter Schouten |
A. Murat Maga and Robin M. D. Beck. 2017. Skeleton of An Unusual, Cat-sized Marsupial Relative (Metatheria: Marsupialiformes) from the middle Eocene (Lutetian: 44-43 million years ago) of Turkey. PLoS ONE. 12(8); e0181712. DOI: 10.1371/journal.pone.0181712
Cenozoic carnivore from Turkey may have evolved without placental competitors phy.so/422094230 via @physorg_com
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