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Sunday, August 22nd, 2021
| Time | Event | ||||||||
| 2:52a | [Herpetology • 2021] Boophis asquithi • An Additional Level of Cryptic Diversity: A New Green-coloured Malagasy Treefrog of the Boophis luteus Species Group (Anura: Mantellidae)
Abstract New genetic, bioacoustic and morphological data on green-coloured Boophis treefrogs from eastern Madagascar reveal an additional level of cryptic diversity in these frogs. Two candidate species, Boophis sp. Ca36 and Ca37, are closely related to each other and to B. sandrae, with uncorrected pairwise distances in the mitochondrial 16S rRNA gene as low as 2.2% between some individuals. However, the three lineages show full concordance between differentiation in the 16S and the nuclear-encoded SACS gene, despite confirmed syntopy of B. sandrae and B. sp. Ca37 in the Ranomafana region, and probable syntopy of B. sp. Ca36 and B. sp. Ca37 in the Andasibe region. Most likely, these lineages are also divergent in advertisement calls, but the available recordings cannot be reliably assigned to either of them. Based on new material collected from various new sites, we here formally name B. sp. Ca36 as new species Boophis asquithi sp. n., and suggest targeted fieldwork on calls and larval stages to allow for a complete and fully conclusive taxonomic revision of this species complex. The example of these frogs illustrates how continued underestimation of cryptic diversity in anurans can lead to incorrect assignment of specimens, and leads us to emphasize the importance of designating as name-bearing types (holotypes) of anurans only individuals whose identity is unambiguous by genetic data or, at least, call recordings reliably assignable to the type specimen. Key words. Amphibia, Anura, Mantellidae, Boophis luteus species group, Boophis elenae, Boophis sandrae, Boophis asquithi sp. n., Madagascar, cryptic species. 
  Boophis asquithi sp. n. Etymology. The specific name is dedicated to Mr. John David Asquith, in recognition of his support of biodiversity research and nature conservation through the BIOPAT initiative. Natural history. Calls probably emitted by B. asquithi were sometimes heard from higher positions in trees around Andasibe in the rainy season, but unfortunately, call recordings are not available of any voucher specimen. The genetically unconfirmed male ZFMK 60027 was calling on 15 January 1995 (at night) from vegetation ca 3 m above the ground. The vocal sac was relatively large, but did not inflate during the vocalization, suggesting that the vocalizations were possibly produced both during expiration and inspiration. The couple from near Tarzanville (Fig. 8) was found in axillary amplexus along a small river in the evening of 12 April 2010. The female had numerous yellowish eggs in its body cavity. A second couple in axillary amplexus probably assignable to this species was found inactive (sleeping) during the day along the road. ZSM 130/2016 was sitting at night on a leaf in the vegetation along the road, less than 1 m above the ground and did not vocalize. Frank Glaw, Jörn Köhler, Angelica Crottini, Philip-Sebastian Gehring, David Prötzel, Lalaina Randriamanana, Franco Andreone and Miguel Vences. 2021. An Additional Level of Cryptic Diversity: A New Green-coloured Malagasy Treefrog of the Boophis luteus Species Group. Salamandra. 57(3); 295-308.  | ||||||||
| 2:53a | [Herpetology • 2021] Phrynoglossus swanbornorum • A New Species of Phrynoglossus Peters, 1867 (Anura: Dicroglossidae) from southeastern Bangladesh, with Comments on the Genera Occidozyga and Phrynoglossus
Abstract We describe a new cryptic species of Phrynoglossus from Chattogram Division, Bangladesh based on an integrative taxonomic analysis based on morphology, phylogenetics, and bioacoustics which unambiguously support the placement of the species in the genus Phrynoglossus. We also present a compilation of published morphological characters for all twelve Phrynoglossus species and two species of Occidozyga as well as comments on taxonomy, morphology, and geographic distribution for the two genera. The new species is found to be most morphologically similar to P. martensii, however a provided set of character states visibly differentiates these two species. Finally, habitat for Phrynoglossus swanbornorum sp. nov. is highly fragmented and faces imminent threats from development and agriculture, and although it is confirmed to occur within government protected areas in the southeastern region of Bangladesh, few wildlife regulations are enforced within them. Thus, following IUCN criteria, we consider the new species as Endangered based on criteria B1ab(i,ii,iii,iv) + 2ab(i,ii,iii,iv). 
Phrynoglossus swanbornorum sp. nov. Recommended vernacular name. English: Swanborn’s Puddle Frog. Bangla: “Chattgai ar gata bang,” which translates to English as “puddle frog from Chattogram.” Generic placement. The new species is assigned to the genus Phrynoglossus based on the following combination of shared adult characters: lacking vomerine teeth and exhibiting a stocky habitus, flattened snout, dorsum or flanks with scattered tubercles, short arms, nuptial pads, horizontal pupils, bony style, forked omosternum, pigmented eggs, distinct or indistinct supratympanic fold, reduced to absent metacarpal webbing, moderate to extensive metatarsal webbing, an elongated inner metatarsal tubercle, indistinct and small tympanum, feebly to moderately developed toes discs, SVL between 15 mm and 61.6 mm in length, and females are larger than males. These adult characters agree with previous descriptions of the genus (see discussion; Taylor, 1962; Iskandar, 1998; Sailo et al., 2009; Inger et al., 2017; Poyarkov et al., 2020; Köhler et al., 2021). Larval characters are unknown. Diagnosis. The new species presents the following characteristics: (1) relatively small sized Phrynoglossus (adult males = 23.15–28.56 mm SVL, adult female = 30.56 mm); (2) stocky habitus; (3) short arms relative to body size (FLL/SVL 0.19%; range 0.18−0.20%, n = 5); (4) head wider than long (HL/HW 0.73%; range 0.69−0.77%, n = 5); (5) snout slightly projecting, triangular in ventral aspect, smaller than horizontal diameter of eye, and flat in lateral aspect; (6) nostril closer to tip of snout than to eye (NS 0.48−0.51 mm, EN 0.92–109 mm); (7) brown dorsum transitioning to brownish grey on flanks with black speckling concentrated around scattered tubercles, nares, mouth, and above eyes; (8) venter uniform cream white, becoming brown with white mottling and groups of minute dark-grey flecks present in the gular region; (9) tuberculate dorsum; (10) laterally oriented eyes and nares; (11) horizontal pupils; (12) vomerine teeth absent; (13) tongue rounded without notch behind; (14) vocal sac single, median, internal, and subgular in males; (15) canthus rostralis rounded; (16) tympanum indistinct, small, and rounded; (17) supratympanic fold distinct and transverses interorbitally at the posterior edge of the eye; (18) fingertips rounded without discs; (19) nuptial pads present in males; (20) palmar tubercles present; (21) metacarpal webbing absent; (22) toe discs feebly developed; (23) inner metatarsal tubercle elongated and compressed, present at base of 1st toe; (24) fringe of skin on outer side of fifth metatarsal absent; (25) tarsal tubercle absent; (26) tarsal fold absent; (27) metatarsal webbing moderate; and (28) lateral line absent. Details of these characteristics are provided in Table 3. Etymology. The specific patronym epithet “swanbornorum” is in genitive plural and refers to members of the Swanborn family, who are generous supporters of the conservation efforts of the Creative Conservation Alliance. Distribution. Phrynoglossus swanbornorum sp. nov. is currently only confirmed to occur within the Chunati Wildlife Sanctuary (21.937533 N, 92.063010 E), Chattogram District at an elevation of ca. 33 m and Teknaf Wildlife Sanctuary (20.887020 N, 92.298954 E), Cox’s Bazar District at an elevation of ca. 26 m (Fig. 1). Natural history: Phrynoglossus swanbornorum sp. nov. is only known to inhabit lowland, coastal, mixed-evergreen forests of southeast Bangladesh, which are bounded to the east by the mountainous Chattogram Hill Tracts. Within this habitat, Phrynoglossus swanbornorum sp. nov. has only been observed near lentic water bodies under dense forest canopy and was notably absent from surrounding agricultural fields and degraded forest. An image of P. swanbornorum sp. nov. in amplexus (see Distribution; Ahm, 2014) demonstrates that this species exhibits inguinal amplexus. Additionally, eggs from the single collected female were pigmented.
Conclusions: We provide morphological, molecular, and bioacoustic evidence that support the validity of the new species, Phrynoglossus swanbornorum sp. nov. This species is currently only confirmed from a relatively small area in southeastern Bangladesh and is considered to have a high risk of extinction due to conspicuous and ongoing threats to the habitat it relies on. We also review and discuss inconsistencies in the literature regarding the genera Phrynoglossus and Occidozyga, particularly that of morphological characters and provide more accurate generic diagnoses for both. Although multiple species of Phrynoglossus and Occidozyga are generally perceived to be widespread and common, this study adds to a growing body of literature demonstrating the existence of undescribed diversity within the genus, thereby masking the true extinction risk of these species (Mulcahy et al., 2018). We suggest further research be conducted to either rectify incongruent descriptions of diagnostic characters and/or better understand intraspecific variation within the genus as well as including additional nuclear genetic data. Doing so could result in further refinement of the provided generic diagnoses. We also suggest that further surveys be conducted in the northern part of Chattogram Hill Tracts, Bangladesh e.g., Pablakhali Wildlife Sanctuary and Kassalong Reserve Forest, and throughout Rakhine State, Myanmar to better understand the distribution of Phrynoglossus swanbornorum sp. nov. Scott Trageser, Hassan Al-Razi, Marjan Maria, Fahimuzzaman Nobel, Md. Asaduzzaman and Shahriar Caesar Rahman. 2021. A New Species of Phrynoglossus Peters, 1867; Dicroglossidae) from southeastern Bangladesh, with Comments on the Genera Occidozyga and Phrynoglossus. PeerJ. 9:e11998. DOI: 10.7717/peerj.11998 | ||||||||
| 3:12a | [Botany • 2021] Three New Species of Polyalthia (Annonaceae) from Thailand and Laos: Polyalthia chalermglinii, P. chantaranothaii & P. chayamaritana
Abstract Three new species of Polyalthia are described and illustrated. Two of the new species are part of the Polyalthia evecta species complex, while the third species belongs to a species group including P. cinnamomea and P. stenopetala. Keywords: Magnoliids, endemism, flora of Thailand, flora of Southeast Asia, Miliuseae
Polyalthia chalermglinii P.Bunchalee & D.M.Johnson, sp. nov. Ecology:— Endemic to peninsular Thailand in tropical rain forest at 950–1000 m. Etymology:—Named in honour of Piya Chalermglin, Thai botanist, horticulturalist and expert on Annonaceae, who assisted in the discovery of this species. Local name:—Thai: tong laeng khao luang (ต้องแล่งเขาหลวง).  Polyalthia chantaranothaii P.Bunchalee & Chalermglin, sp. nov. Ecology:—Scattered along streams and waterfalls in mixed deciduous forest or dry evergreen forest at 200–850 m. Etymology:—In honour of Pranom Chantaranothai, Thai professor of botany and benefactor of many plant taxonomists, who assisted the first author in analyzing the morphology of this species. Local name:—Thai: tong laeng doi (ต้องแล่งดอย); nom noi (นมน้อย) (Phitsanulok). Polyalthia chayamaritana P.Bunchalee & N.A.Murray, sp. nov. Ecology:—Known only from moist mountain slopes in dry evergreen forest at 250–375 m. Etymology:—In honour of Kongkanda Chayamarit, expert on the Thai flora and executive editor of the Flora of Thailand project. Local name:—Thai: thon ammarit (ต้นอัมฤทธิ์). Pasakorn Bunchalee, David M. Johnson, Nancy A. Murray and Piya Chalermglin. 2021. Three New Species of Polyalthia (Annonaceae) from Thailand and Laos. Phytotaxa. 512(4); 272–282. DOI: 10.11646/phytotaxa.512.4.3 | ||||||||
| 3:16a | [Botany • 2021] Magnolia zotictla (Magnoliaceae: sect. Macrophylla) • A New Species from the southern Sierra Madre Oriental, México
Abstract A new species of Magnolia from the southern Sierra Madre Oriental, Mexico, is described and illustrated, providing information about its habitat distribution, ecology, biogeography and conservation status. After 12 fieldwork expeditions near the border of the states of Hidalgo and Puebla, we have developed morphological, ecological and biogeographic data to support recognition of populations from Acaxochitlán, Hidalgo and Pahuatlán, Puebla as a distinct species of Magnolia sect. Macrophylla. A key to species of this section and a distribution map for Mexican taxa are provided. The species was assessed as critically endangered (CR). Keywords: Cloud-forest species, Mexican endemic, Mexican flora, Magnolia dealbata Magnolia zotictla A.Sánch.-Gonz., Gut.-Lozano & A.Vázquez, spec. nov. Type:—MEXICO. Hidalgo: Municipality Acaxochitlán, Zotictla, 0.3 km al SE de San Miguel del Resgate, bosque mesófilo de montaña, 1743 m, 20°13’32.5”N, 98°09’48”W, 5 May 2021 (fl & fr) Gutiérrez-Lozano et al. 10186 (holotype: HGOM; isotypes: ENCB, IBUG, OAX). Magnolia zotictla is morphologically more similar to the distant M. dealbata than to nearby M. rzedowskiana. It shares with the former habit size and large, deciduous, abaxially glaucous leaves. However, it differs from M. dealbata in having pubescent pedicels (vs. glabrescent), flowers with sepals erect (uncurled) at female phase (vs. curling outwards with petals maintaining a compact oblong conic shape), most petals strongly curled outward (vs. occasionally curling), stamens 1.3–1.8 (vs. 1.9–2.1 cm long), stigmas crowded, strongly curved outward and covering the carpels (vs. loose, slightly curved outward and exposing the carpels), young fruits velvety (vs. glabrescent), mature fruits pinkish-purplish to dark red (vs. dark greenish to yellowish), carpels densely pubescent (vs. glabrescent) and carpel beak acuminate to caudate with the stigmatic tip sometimes persistent during dehiscence (vs. acute to blunt with a deciduous stigmatic tip). Ethnobotany:—In the ancient cultures of America and Asia, magnolias were known for their ornamental and pharmacological use, essentially to alleviate various types of human pathologies (Pfaffman 1975, Dodd 1980, Gutiérrez-Carvajal 1993, Gutiérrez and Vovides 1997, Satyajiy et al. 2002, Vázquez-García et al. 2015). This species is known locally as magnolia, eloxóchitl or yeloxóchitl (cornflower in Náhuatl), and its showy and fragrant flowers are usually collected for sale locally. The most important use in the region is to prepare floral infusions to treat heart problems, nerve problems and scars. It is also used as an ornamental plant. A few people from Santa Catarina, Toxtla and Zotictla sell the petals largely on Sunday in the local markets of Hidalgo (Actopan, Pachuca, San Miguel Resgate and Tulancingo), Puebla (Honey, Pahuatlán), Veracruz (Poza Rica and Tuxpan), and Mexico City (Sonora traditional market). A kg of petals sells at $4–6 USD for use as infusions. Arturo Sánchez-González, Marisol Gutiérrez-Lozano, Reyna Domínguez Yescas, Adriana Gisela Hernández-Álvarez, A. Salomé Ortega-Peña and J. Antonio Vázquez-García. 2021. Magnolia zotictla (Magnolia sect. Macrophylla, Magnoliaceae): A New Species from the southern Sierra Madre Oriental, México. Phytotaxa. 513(4); 271-281. DOI: 10.11646/phytotaxa.513.4.1 Resumen: Se describe e ilustra una nueva especie de Magnolia del sur de la Sierra Madre Oriental, México, proporcionando información sobre su distribución, hábitat, ecología, biogeografía y estado de conservación de la especie. A través de 12 expediciones de trabajo de campo cerca de la frontera de los estados de Hidalgo y Puebla, se recabaron datos morfológicos, ecológicos y biogeográficos que apoyan el reconocimiento de las poblaciones de Acaxochitlán, Hidalgo y Pahuatlán, Puebla como una especie distintiva de Magnolia de la sección Macrophylla. Se proporciona una clave de las especies de esta sección y un mapa de distribución de los taxones mexicanos. La especie fue evaluada como en peligro crítico (CR). Palabras clave: flora del bosques nuboso, Especie endémica mexicana, flora mexicana, Magnolia dealbata | ||||||||
| 1:22p | [Paleontology • 2021] New Fossils of Jurassic Ophiurid Brittle Stars (Ophiuroidea; Ophiurida) provide evidence for early Clade Evolution in the Deep Sea; Ophiogojira labadiei, Ophiogojira andreui & Ophioduplantiera noctiluca
Abstract Understanding of the evolutionary history of the ophiuroids, or brittle stars, is hampered by a patchy knowledge of the fossil record. Especially, the stem members of the living clades are poorly known, resulting in blurry concepts of the early clade evolution and imprecise estimates of divergence ages. Here, we describe new ophiuroid fossil from the Lower Jurassic of France, Luxembourg and Austria and introduce the new taxa Ophiogojira labadiei gen. et sp. nov. from lower Pliensbachian shallow sublittoral deposits, Ophiogojira andreui gen. et sp. nov. from lower Toarcian shallow sublittoral deposits and Ophioduplantiera noctiluca gen. et sp. nov. from late Sinemurian to lower Pliensbachian bathyal deposits. A Bayesian morphological phylogenetic analysis shows that Ophiogojira holds a basal position within the order Ophiurida, whereas Ophioduplantiera has a more crownward position within the ophiurid family Ophiuridae. The position of Ophioduplantiera in the evolutionary tree suggests that family-level divergences within the Ophiurida must have occurred before the late Sinemurian, and that ancient slope environments played an important role in fostering early clade evolution. Keywords: Echinodermata, ophiuroids, deep sea, phylogeny, evolution  Systematic palaeontology Class Ophiuroidea Gray, 1840 Subclass Myophiuroidea Matsumoto, 1915 Infraclass Metophiurida Matsumoto, 1913 (crown-group of Ophiuroidea) Superorder Euryophiurida O'Hara et al. (2017) Order Ophiurida Müller & Troschel (1840) Suborder Ophiurina Müller & Troschel, 1840 sensu O'Hara et al. [5] Family unknown Genus Ophiogojira gen. nov. Etymology: Genus named in honour of French metal band Gojira, for producing songs of an unfathomable intensity, beautifully dark and heavy, and exploring the abyss of life and death, of human strength and error, and of thriving and yet threatened oceans. Ophiogojira labadiei sp. nov. Etymology: Species named after Jean-Michel Labadie, bass player of French metal band Gojira. Holotype: MnhnL OPH159 Type locality and stratum: succession of clays and thin lumachellic beds with siderite nodules, dated to the late Early Pliensbachian (Davoei chronozone, Early Jurassic), near Sedan, Ardennes, France. Ophiogojira andreui sp. nov. Etymology: Species named after Christian Andreu, guitar player of French metal band Gojira. Holotype: MnhnL OPH167 Type locality and stratum: dark grey marls, Ottempt Member of the Aubange Formation, earliest Toarcian (Tenuicostatum chronozone, Early Jurassic), Uerschterhaff near Sanem, Luxembourg. Family Ophiuridae Müller & Troschel, 1840 Genus Ophioduplantiera gen. nov. Etymology: Genus named after Joseph (Joe) and Mario Duplantier of French metal band Gojira, to honour their inspiring artistic achievements and their authentic, compassionate personalities. Ophioduplantiera noctiluca sp. nov. Etymology: Species name formed from Latin ‘noctiluca’, literally translating into ‘night light’, in reference to the bioluminescent dinoflagellate genus Noctiluca, commonly called ‘sea sparkle’, because the name-bearing ophiuroid species brings light into the dark origins of the Ophiuridae. Holotype: MnhnL OPH170 Type locality and stratum: marls from the Kehlbach and Scheck Members within the Adnet Formation, late Sinemurian to late Pliensbachian (Raricostatum to Margaritatus chronozones, Early Jurassic), Glasenbach Gorge near Salzburg, Austria.  Ben Thuy, Lea D. Numberger-Thuy and Tania Pineda-Enríquez. 2021. New Fossils of Jurassic Ophiurid Brittle Stars (Ophiuroidea; Ophiurida) provide evidence for early Clade Evolution in the Deep Sea. Proceedings of the Royal Society B. DOI: 10.1098/rsos.210643  | ||||||||
| 1:41p | [Herpetology • 2021] A Large and Unusually Thick-shelled Turtle Egg (Cryptodira: Nanhsiungchelyidae) with Embryonic Remains from the Upper Cretaceous of China
Abstract Turtle eggs containing embryos are exceedingly rare in the fossil record. Here, we provide the first description and taxonomic identification, to our knowledge, of a fossilized embryonic turtle preserved in an egg, a fossil recovered from the Upper Cretaceous Xiaguan Formation of Henan Province, China. The specimen is attributed to the Nanhsiungchelyidae (Pan-Trionychia), an extinct group of large terrestrial turtles (possibly the species Yuchelys nanyangensis). The egg is rigid, spherical, and is one of the largest and thickest shelled Mesozoic turtle eggs known. Importantly, this specimen allowed identification of other nanhsiungchelyid egg clutches and comparison to those of Adocidae, as Nanhsiungchelyidae and Adocidae form the basal extinct clade Adocusia of the Pan-Trionychia (includes living soft-shelled turtles). Despite the differences in habitat adaptations, nanhsiungchelyids (terrestrial) and adocids (aquatic) shared several reproductive traits, including relatively thick eggshells, medium size clutches and relatively large eggs, which may be primitive for trionychoids (including Adocusia and Carrettochelyidae). The unusually thick calcareous eggshell of nanhsiungchelyids compared to those of all other turtles (including adocids) may be related to a nesting style adaptation to an extremely harsh environment. Keywords: Nanhsiungchelyidae, China, thick egg, shell, Upper Cretaceous, embryo, turtle egg Systematic palaeontology: Order Testudines Linnaeus, 1758 Infraorder Cryptodira Cope, 1868 Superfamily Trionychoidae Fitzinger, 1826 Family Nanhsiungchelyidae Yeh, 1966 Genus and species indet. 
 Material. CUGW EH051, a complete egg containing embryonic remains. Locality and horizon. Specimen CUGW EH051 was collected in Neixiang County, Nanyang City, Henan Province, China (electronic supplementary material, figure S1); Xiaguan Formation, Upper Cretaceous. ....   
 Yuzheng Ke, Rui Wu, Darla K. Zelenitsky, Don Brinkman, Jinfeng Hu, Shukang Zhang, Haishui Jiang and Fenglu Han. 2021. A Large and Unusually Thick-shelled Turtle Egg with Embryonic Remains from the Upper Cretaceous of China. Proceedings of the Royal Society B. DOI: 10.1098/rspb.2021.1239 | ||||||||
| 3:25p | [Herpetology • 2021] Chirixalus pantaiselatan • A New Species of Chirixalus Boulenger, 1893 (Anura: Rhacophoridae) from the Lowland Forests of Java, Indonesia
Abstract The Old World tree frog genus Chirixalus is distributed from northeastern India, southern China, continental Southeast Asia to Sumatra and Java. The species of this genus were previously assigned to the genera Chiromantis or Philautus. Here, we describe a newly discovered species of Chirixalus from Java. Chirixalus pantaiselatan, new species, is a small rhacophorid frog (male snout-vent length = 25.3–28.9 mm) that can be distinguished from all congeners using a combination of morphological, molecular, and advertisement call characteristics. Key words. bioacoustics, Chirixalus, citizen science, taxonomy
Chirixalus pantaiselatan, new species Diagnosis. The new species was assigned to the genus Chirixalus based on the following external characters: horizontal pupil, tongue free and deeply notched, distinct tympanum, opposed fingers, toes webbed, tips of fingers and toes dilated into disks, outer metatarsal separated by web, terminal phalanges obtuse (Boulenger, 1893). The new species can be distinguished from its congeners by the combination of the following characters: snout truncate; SVL 25.3–28.9 mm in males; uniform lateral body colouration, without a light dorsolateral stripe from canthus rostralis through upper orbital to the body or a light lateral stripe from under orbital to mid-body; brown dorsal colouration, with an indistinct transverse dark brown patch on intercanthal and upper eyelid to interorbital area continued by an indistinct irregular hexagon-like shape covering the parietoscapular region, and a diamond-like shape on suprascapular region to mid-body; distinct dark brown tympanum; third finger disk diameter about half that of tympanum and unique advertisement call consisting of short (2–9 notes), medium (11–17 notes), and long calls (22–46 notes). Calls are 185.71–1,644.28, 1,420.78–2,708.46, and 3,292.29–7,055.79 ms in duration, respectively, with note durations of 4.14–18.03, 4.82–15.19, and 4.78–24.88 ms, respectively. Dominant frequencies ranging from 2.34–3.94, 2.53–3.75, and 2.72–3.94 kHz for calls of short, medium, and long duration. Etymology. The specific name is an Indonesian adjective incorporating the words “Pantai,” meaning “coast,” and “Selatan,” meaning “south.” This name refers to the locality of this species in southern Java. Indonesian Name. Katak-pucat pantaiselatan Misbahul Munir, Amir Hamidy, Mirza Dikari Kusrini, Umar Fhadli Kennedi, Mohammad Ali Ridha, Dzikri Ibnul Qayyim, Rizky Rafsanzani and Kanto Nishikawa. 2021. A New Species of Chirixalus Boulenger, 1893 (Anura: Rhacophoridae) from the Lowland Forests of Java. RAFFLES BULLETIN OF ZOOLOGY. 69; 219–234 |
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