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Sunday, April 24th, 2022
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2:05p |
[Ichthyology • 2019] Astyanax varii • A New Species of Astyanax (Characiformes: Characidae) from the rio de Contas basin, Bahia, Brazil
 | Astyanax varii
Zanata, Burger, Vita & Camelier, 2019
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ABSTRACT A new species of Astyanax from tributaries of the rio de Contas, Bahia, Brazil, is described. The new species differs from congeners by having three horizontal series of scales from lateral line to pelvic-fin origin and the distal margin of third infraorbital distinctly separated from vertical and horizontal limbs of preopercle, leaving a broad area not covered by superficial bones. The new species further differs from most congeners by the presence of bony hooks on all fins of mature males. Particularly from congeners occurring in rivers of the Northeastern Mata Atlântica freshwater ecoregion, it further differs by having the highest body depth just anterior to the dorsal-fin origin, 34-37 pored lateral line scales, a vertically elongated conspicuous dark humeral blotch reaching below the lateral line and a conspicuous dark wide midlateral stripe extending from the clear area on the rear of the humeral blotch to the end of middle caudal-fin rays and forming an inconspicuous blotch on caudal peduncle.
Keywords: Endemism; Northeastern Mata Atlântica freshwater ecoregion; Sexual dimorphism; Taxonomy
 | Astyanax varii, Brazil, Bahia State: (a) MZUSP 121062, holotype, 41.4 mm SL,Ubaitaba, lower rio de Contas basin, rio Coricó; (b) live paratype, not catalogued. |
Astyanax varii, new species
Astyanax sp. 6.-Camelier, Zanata (2014):686, tab. 1 [listed; species from the Northeastern Mata Atlântica freshwater ecoregion]. Astyanax sp.-Barreto et al. (2018): 1158 [citation; co-occurence with Nematocharax varii].
Diagnosis. Astyanax varii can be distinguished from most congeners, except A. brachypterygium Bertaco & Malabarba, A. brucutu, A. cremnobates Bertaco & Malabarba, A. epiagos, A. eremus Ingenito & Duboc, A. gymnogenys Eigenmann, A. rupestris Zanata, Burger & Camelier, A. taeniatus (Jenyns), A. totae Haluch & Abilhoa, and A. varzeae Abilhoa & Duboc, by having the distal margin of third infraorbital distinctly separated from vertical and horizontal limbs of preopercle (Fig. 2), leaving a broad area without superficial bones (vs. margins of the third infraorbital close to the limbs of preopercle, with narrow or no space between the two bones). The new species differs from the aforementioned species by having three horizontal series of scales from the lateral line to the pelvic-fin origin (vs. four or more series of scales) and by having small bony hooks on all fins of mature males (vs. bony hooks absent or not present on all fins of mature males). Astyanax varii also differs from the species listed above by having highest body depth approximately at vertical through dorsal-fin origin (vs. body deepest on a vertical through middle or posterior portion of pectoral fin in A. brachypterygium, A. cremnobates, A. epiagos, A. eremus, A. gymnogenys, A. rupestris, A. totae, and A. varzeae), five horizontal series of scales from the dorsal-fin origin to the lateral line (vs. six or more in A. brachypterygium, A. brucutu, A. cremnobates, A. eremus, A. rupestris, A. totae, A. taeniatus, and A. varzeae), 14 horizontal scale rows around caudal peduncle (vs. 15 or more in A. brachypterygium, A. eremus, A. gymnogenys, and A. totae), and 34-37 pored lateral line scales (vs. 38 or more in A. eremus, A. gymnogenys, and A. taeniatus).
Etymology. The specific name varii is in honor to the ichthyologist Richard P. Vari for his friendship, mentoring, and outstanding contribution to the systematic of South American freshwater fishes.
Angela Maria Zanata, Rafael Burger, eorge Vita and Priscila Camelier. 2019. A New Species of Astyanax (Characiformes: Characidae) from the rio de Contas basin, Bahia, Brazil. Neotropical Ichthyology. 17(3) DOI: 10.1590/1982-0224-20190061
RESUMO: Uma espécie nova de Astyanax de tributários do rio de Contas, Bahia, Brasil, é descrita. A espécie nova difere das congêneres por ter três séries horizontais de escamas entre a linha lateral e a origem da nadadeira pélvica e margem distal do terceiro infraorbital distintamente separada dos ramos vertical e horizontal do pré-opérculo, deixando uma ampla área sem ossos superficiais. A espécie nova difere, ainda, da maioria das congêneres pela presença de ganchos ósseos em todas as nadadeiras de machos maduros. Particularmente das congêneres que ocorrem nas drenagens da Ecorregião Mata Atlântica Nordeste, difere ainda por ter a maior altura do corpo imediatamente anterior à origem da nadadeira dorsal, 34-37 escamas perfuradas na linha lateral, presença de uma mancha umeral conspícua, verticalmente alongada, ultrapassando ventralmente a linha lateral e uma faixa escura mediana lateral larga, estendendo-se da área clara após a mancha umeral até o fim dos raios medianos caudais e formando uma mancha inconspícua no pedúnculo caudal. Palavras-chave: Dimorfismo sexual; Ecorregião Mata Atlântica Nordeste; Endemismo; Taxonomia
| 2:33p |
[Botany • 2021] Isodon brevipedunculatus (Lamiaceae: Isodoninae) • A New Species of from southern China
 | Isodon brevipedunculatus Y.P. Chen & C.L. Xiang,
in Chen, Huang, Zhao & Xiang, 2021 |
Abstract
Isodon brevipedunculatus, a new species from southern China, is described and illustrated. The phylogenetic position of the new species within the genus was analyzed based on two nuclear ribosomal DNA regions and an ingroup sampling of about 80% of Asian species of Isodon. The results show that I. brevipedunculatus is recovered in a clade that consists of species mainly with glandular mericarps and that are distributed in the Sino-Japanese region. Combining molecular and geographical evidence, our study reveals that I. brevipedunculatus is most closely related to Isodon amethystoides and Isodon bifidocalyx, but differs from the former in lamina shape, number of flowers per cyme, and peduncle length, and from the latter in lamina indumentum, calyx morphology, and corolla length.
Keywords: Himalaya-Hengduan Mountains region, Isodoninae, Mericarp, Sino-Japanese region
 | Isodon brevipedunculatus Y.P. Chen & C.L. Xiang (A) habit; (B) flower; (C) dissected calyx; (D) dissected corolla; (E) pistil; (F) mericarp.
Drawn by L. Wang. |
 | Isodon brevipedunculatus Y.P. Chen & C.L. Xiang (from the type locality). (A) habit; (B) leaves; (C, D) panicles; (E) inflorescence; (F, G) infructescences; (H, I) flowers; (J) mericarps.
Photographed by Y.P. Chen. |
Isodon brevipedunculatus Y.P. Chen & C.L. Xiang, sp. nov.
Diagnosis: The new species is morphologically similar to Isodon amethystoides and I. bifidocalyx, but differs from the former in its broadly ovate to ovate laminae (vs. ovate to lanceolate), 3–7-flowered cymes (vs. 7–15-flowered), and 1–2 mm long peduncles (vs. 1–4 cm long), and differs from the latter in having densely pubescent laminae (vs. subglabrous), flowering calyces slightly 2-lipped to 1/3 of their length with teeth subequal (vs. strongly 2-lipped to 1/2 their length with teeth unequal), erect fruiting calyces (vs. declinate), and corollas ca. 4 mm long (vs. ca. 8 mm long).
Distribution and habitat: Isodon brevipedunculatus is currently known from Guangdong Province and Hunan Province in southern China (Fig. 4). It can be found in forests or on grassy slopes at altitudes of 600–1250 m.
Etymology: The specific epithet refers to the short peduncles of the new species, as compared to one of the most similar species, Isodon amethystoides.
Chinese name: Duan Geng Xiang Cha Cai (短梗香茶菜).
Ya-Ping Chen, Cun-Zhong Huang, Yue Zhao and Chun-Lei Xiang. 2021. Molecular and Morphological Evidence for A New Species of Isodon (Lamiaceae) from southern China. Plant Diversity. 43(1); 54-62. DOI: 10.1016/j.pld.2020.06.004
| 2:35p |
[Botany • 2022] Impatiens godfreyi (Balsaminaceae) • A New Species from the southern Western Ghats, India
 | Impatiens godfreyi P.S.S. Richard & Karupp.
in Richard, Karuppusamy & Ravichandran, 2022. |
Abstract Impatiens godfreyi, a new species from mid-elevation evergreen forests of Kalakkad Mundanthurai Tiger Reserve, southern Western Ghats, is described and illustrated. The new species is similar to I. disotis Hook.f. and mainly differs in having perennial branched woody subshrub up to 200 cm high, ovate-elliptic leaves, 2–3-flowered umbel inflorescence, orbicular standard petals with large scarlet-red speckled inside, overlapping lateral corolla lobes with mucronate apex, single lobed distal corolla lobe with a rounded apex and obconical lip with a tubular incurved spur. The morphological characters, distribution, phenology, conservation status, photographs, and taxonomical key for the section subumbellatae are provided for easy identification. Keywords: Agasthyamalai Biosphere Reserve, Endemic, Impatiens, New species, Tamil Nadu, Taxonomy
 | Impatiens godfreyi P.S.S. Richard & Karupp.: A, habit: B, leaf; C, inflorescence; D, flower – lateral view; E, flower – front view; F, flower petals – dissected; G, Anther; H, Capsule; I, Seeds. |
 | Impatiens godfreyi P.S.S. Richard & Karupp.: A, habit; B, leaf; C, inflorescence; D, flower – lateral view; E, flower – front view; F, Anther; G, flower petals – dissected; H, Capsule; I, Seeds. |
Impatiens godfreyi P.S.S. Richard and Karupp. sp. nov.
Diagnosis. Impatiens godfreyi is morphologically similar to I. disotis, but it can be easily distinguished by perennial branched, woody habit (vs. annual unbranched herbaceous), ovate-elliptic leaves with 5–8 cm long petiole (vs. ovate-lanceolate leaves with 4 cm petiole), 2–3-flowered umbel inflorescence (vs. 6–8-flowered raceme), orbicular standard petal with large orbicular scarlet-red patch inside and emarginate apex (vs. ovate standard with purplish red patch inside, acute apex), large obovate lateral corolla lobe overlapping on distal one with a rounded tip and mucronate apex (vs. small oblong lateral corolla lobe not overlapping on distal one with obovate tip and acute apex), single lobed distal corolla, rounded at apex, obliquely clefted near the inner apex (vs. two-lobed distal corolla with acute apex), obconical golden yellow lip (vs. boat-shaped pale yellow lip), 12–20 mm long tubular incurved spur (vs. 6–13 mm long dilated spur). The flowers with a mixture of yellow, green, red, and white combination of balsam flowers are quite unique in this group, and therefore, permits easy identification (Table 1).
Eponym: This species is named in honor of Mr. C. Godfrey Webb-Peploe, a Christian missionary, social worker, and naturalist from the Dohnavur Fellowship, Tirunelveli, Tamil Nadu, to recognize his pioneering efforts to study, document, and conserve the mega fauna and flora of Tirunelveli hills, southern Western Ghats. His “Field notes on the mammals of south Tirunelveli, South India,” published in the year 1947, is the earliest record on the fauna of this region. Before publishing the floristic data, Godfrey faced early death in his 30s on 19th February 1949. He also illustrated the same species in his field note.
Paulraj Selva Singh Richard, Subbiah Karuppusamy and Vellingiri Ravichandran. 2022. Impatiens godfreyi (Balsaminaceae), A New Species of Balsam from the southern Western Ghats, India. Journal of Asia-Pacific Biodiversity. 15(1); 138-144. DOI: 10.1016/j.japb.2021.12.001 | 2:35p |
[Botany • 2022] Hemilophia cardiocarpa (Brassicaceae) • A New Species from Yunnan, southwest China
 | Hemilophia cardiocarpa Huan C. Wang, Shao Y. Liu & Z. T. Ren,
in Liu, Ren, Zhao, Hu & Wang, 2022. |
Abstract Hemilophia cardiocarpa (Brassicaceae), the sixth species of the Chinese endemic genus Hemilophia, is described and illustrated. This plant is found in the Jiaozishan Mountains in Dongchuan District, northern Yunnan, southwest China. Morphologically, it shows close relationships with H. rockii and H. pulchella, but differs from it in the leaf shape and size, inflorescence, flower size, shape of fruit and length of its pedicel. An updated key to the taxa of Hemilophia is provided.
Keywords: Alpine flora, Cruciferae, endemism, Hemilophia rockii, Jiaozishan Mountains
 | Hemilophia cardiocarpa Huan C. Wang, Shao Y. Liu & Z. T. Ren. A habit B a portion of stem showing trichomes C cauline leaf D flower (apical view) E petal F median filament G lateral filament H fruit (front view) I fruit (lateral view) J fruit after removal of valves and seeds K seed. |
 | Hemilophia cardiocarpa Huan C. Wang, Shao Y. Liu & Z. T. Ren. A–C habit D a portion of stem with two cauline leaves E inflorescences in apical view F flower G fruit (front view) H fruit (lateral view). |
Hemilophia cardiocarpa Huan C. Wang, Shao Y. Liu & Z. T. Ren, sp. nov.
Diagnosis: Hemilophia cardiocarpa is most similar to H. rockii, but clearly differs from the latter by its cauline leaves 5–8 mm long (vs. 3–6 mm long in H. rockii), 2–4 mm (vs. 1–3 mm) wide, pedicel of fruit shorter than or nearly equal to its leaflike bract (vs. significantly longer than leaflike bract) and fruit cordate (vs. spindle or narrowly oblong) in shape.

Etymology: The specific epithet cardiocarpa is derived from the Greek words “kardio” (heart) and “karpos” (fruit), referring to the fruit shape of this new species.
Vernacular name: Chinese Mandarin: xin guo ban ji ji (心果半脊荠).
Distribution and ecology: Hemilophia cardiocarpa appears to be a rare species endemic to the Jiaozishan Mountains, which are located in the northern Yunnan Province of southwest China with a highest elevation of 4344.1m and near to the Jinsha River. The new species grows on basaltic screes or open slopes at elevations ranging from 3900 to 4300 m, its association mainly include Arenaria weissiana Hand-Mazz. (Caryophyllaceae), Draba amplexicaulis Franch. (brassicaceaee), Scrophularia delavayi Franch.(Scrophulariaceae), Meconopsis integrifolia (Maxim.) Franch. (Papaveraceae), Ranunculus hirtellus var. orientalis W. T. Wang (Ranunculaceae) and Veronica szechuanica subsp. sikkimensis (Hook.f.) Hong (Plantaginaceae).
Shao-Yun Liu, Zheng-Tao Ren, Chang-You Zhao, Chun-Xiang Hu and Huan-Chong Wang. 2022. Hemilophia cardiocarpa (Brassicaceae), A New Species from Yunnan, southwest China. PhytoKeys. 194: 95-103. DOI: 10.3897/phytokeys.194.82737
| 2:35p |
[Ichthyology • 2022] Revalidation of the Genus Ichthyocoris Bonaparte, 1840 (Blenniiformes: Blenniidae)
 | (A) Ichthyocoris atlantica, Morocco, Ouerrha River; (B) I. economidisi, Greece, Lake Trichonis; (C) I. fluviatilis, Spain, Jerea River (Ebro drainage).
in Duquenne-Delobel, Doadrio & Denys, 2022. (photos: I. Doadrio & R. Covain) |
Abstract Combtooth blennies belonging to the genus Salaria were known to have marine and freshwater species. However, recent molecular studies highlighted this genus as paraphyletic, clearly distinguishing both marine and freshwater species. In this paper, we revalidate the genus Ichthyocoris, which corresponds to the freshwater species: Ichthyocoris atlantica (Doadrio, Perea et Yahyaoui, 2011), new combination, Ichthyocoris economidisi (Kottelat, 2004), new combination, and Ichthyocoris fluviatilis (Asso y del Rio, 1801), new combination. It is distinguishable by the presence of brownish bars on the flanks not contrasted with black dots conferring a marble coat, a dorsal fin slightly notched between spined and soft rays (except for I. atlantica), 16–18 dorsal-fin soft rays, 16–20 anal-fin soft rays, 34–38 vertebrae, 8–9 circumorbital pores, 8–11 preopercular pores, and 3 supratemporal pores. The genus Salaria corresponds to the marine species Salaria basilisca (Valenciennes, 1836) and Salaria pavo (Risso, 1810).
Keywords: combtooth blennies, generic concept, integrative taxonomy, Salaria
 | Lateral view of Ichthyocoris spp.: I. atlantica, MNCN 280135, 61 mm SL, Morocco, Ouerrha River (Sebou drainage) at Ouazzane
(A; photo credits: I. Doadrio); I. economidisi, MHNG 2641.89, holotype, 60.8 mm SL, Greece, Lake Trichonis east of Panetolio
(B; photo credits: R. Covain); I. fluviatilis, 89 mm SL, Spain, Jerea River (Ebro drainage) at Virués
(C; photo credits: I. Doadrio). |
Family Blenniidae
Ichthyocoris Bonaparte, 1840 Type species: Salarias varus Risso, 1827.
Synonyms: Salariopsis Vecchioni, Ching, Marrone, Arculeo, Hundt et Simons, 2022
Included species: Three species: Ichthyocoris atlantica (Doadrio, Perea et Yahyaoui, 2011), new combination; Ichthyocoris economidisi (Kottelat, 2004), new combination; Ichthyocoris fluviatilis (Asso y del Rio, 1801), new combination.
Diagnosis: Ichthyocoris is distinguishable from Salaria by the presence of brownish bars on the flanks not contrasted with black dots conferring a marble coat (Fig. 1) (vs. brownish bars on the flanks very contrasted with blue stripes and dots conferring a marbled coat; Fig. 2); dorsal-fin slightly notched between spined and soft rays (Fig. 1) except for I. atlantica (vs. not notched; Fig. 2); 16–18 dorsal-fin soft rays (vs. 21–27); 16–20 anal-fin soft rays (vs. 20–28); 34–38 vertebrae (vs. 38–44); 8–9 circumorbital pores (vs. 6–7); 8–11 preopercular pores (vs. 6–8); 3 supratemporal pores (vs. 2) (Table 1).
Distribution: Ichthyocoris is present in drainages of the Mediterranean basin, in catchments of the Atlantic coast in Morocco and Spain as well as in the Black Sea.
Emma Duquenne-Delobel, Ignacio Doadrio and Gaël P. J. Denys. 2022. Revalidation of the Genus Ichthyocoris Bonaparte, 1840 (Actinopterygii: Blenniiformes: Blenniidae). Acta Ichthyologica et Piscatoria. 52(1): 35-41. DOI: 10.3897/aiep.52.79458
| 2:37p |
[Ichthyology • 2022] Haplochromis aquila, H. kimondo, H. rex, H. simba, etc. • From A Pair to A Dozen: The Piscivorous Species of Haplochromis (Cichlidae) from the Lake Edward System
 | Haplochromis rex, H. aquila,
Haplochromis simba, H. kimondo, Haplochromis glaucus, H. falcatus,
Vranken, Van Steenberge, Heylen, Decru & Snoeks, 2022
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ABSTRACT Piscivory is a common trophic niche among cichlids of the East African Great Lakes, including Lakes Edward and George. From these two lakes, we examined the taxonomic diversity of cichlid species with a piscivorous morphology. Prior to this study, two piscivorous species were formally described, Haplochromis squamipinnis and H. mentatus. We redescribe both species and describe an additional ten new species of Haplochromis with a piscivorous morphology: H. latifrons sp. nov., H. rex sp. nov., H. simba sp. nov., H. glaucus sp. nov., H. aquila sp. nov., H. kimondo sp. nov., H. falcatus sp. nov., H. curvidens sp. nov., H. pardus sp. nov., and H. quasimodo sp. nov. All twelve species differ in dominant male colour pattern (unknown for H. latifrons sp. nov. and H. curvidens sp. nov.) and morphological traits. The species can be divided into two morphological groups: the macrodontic piscivores and the microdontic piscivores. This division potentially reflects an ecological differentiation in habitat use, hunting technique, prey species, and prey size. We conclude that some 12–20% of the species from the cichlid assemblage of Lake Edward have a piscivorous morphology.
Keywords: Adaptive radiation, haplochromines, Harpagochromis, Prognathochromis, new species
Phylum Chordata Haeckel, 1874 Class Actinopterygii Klein, 1885 Order Cichliformes Betancur-R et al., 2013
Family Cichlidae Bonaparte, 1840 Subfamily Pseudocrenilabrinae Fowler, 1934 Tribe Haplochromini Poll, 1986
Genus Haplochromis Hilgendorf, 1888 Haplochromis Hilgendorf, 1888: 76 (as a subgenus of Chromis Cuvier, 1814).
Haplochromis mentatus Regan, 1925
Etymology: Specific name not explained in original description, probably derived from the Latin ‘mentum’ for ‘chin’; probably referring to the protruding lower jaw (i.e., projecting lower jaw sensu Regan 1925).
Haplochromis squamipinnis Regan, 1921
Etymology: Specific name not explained in original description, from the Latin ‘squamus’ for ‘scale’, and ‘pinnis’ for ‘fin’; probably referring to minute scales on basal parts of dorsal and anal fins.
Haplochromis latifrons sp. nov.
Etymology: Specific name from Latin ‘latus’ for ‘wide’ and ‘frons’ for ‘forehead’; referring to very broad interorbital area for a piscivorous species.
Haplochromis rex sp. nov.
Etymology: Specific name from the Latin ‘rex’ for ‘king’ (one that holds a preeminent position); referring to very small eyes, deep cheeks, and strong jaws set with large and acute teeth indicating this piscivore has most specialised morphology among all piscivores from the Lake Edward system to hunt on large prey (Barel et al. 1977).
Haplochromis simba sp. nov.
Etymology: Specific name from Swahili ‘simba’ for ‘lion’; referring to yellow body, orange cheeks that resemble manes, and predatory morphology.
Haplochromis glaucus sp. nov.
Etymology: Specific name from the Latin ‘glaucus’ for ‘greyish blue’; referring to grey and light-blue colour pattern of all adult specimens.
Haplochromis aquila sp. nov.
Etymology: Specific name from the Latin ‘aquila’ for ‘eagle’; referring to predatory morphology and large eyes.
Haplochromis kimondo sp. nov.
Etymology: Specific name from the Swahili ‘kimondo’ for ‘meteor’; referring to blunt head, pyriform body with mid-lateral band, and yellow colouration of ventral part of body.
Haplochromis falcatus sp. nov.
Etymology: Specific name from the Latin ‘falcatus’ for ‘sickle-shaped’; referring to acutely pointed sickle-like outer oral teeth.
 | Haplochromis rex sp. nov. a. c–d. Photographs of freshly caught specimens. c. Holotype, a dominant male. d. A female (RMCA 2017.006.P.0355; 135.7 mm SL) to illustrate the live colour patterns. Haplochromis simba sp. nov. c–d. Photographs of freshly caught specimens. c. Dominant male (RMCA 2016.035.P.0224; 97.9 mm SL). d. Female (RMCA 2018.008.P.0348; 109.0 mm SL) to illustrate the live colour patterns. Haplochromis glaucus sp. nov. c–d. Photographs of freshly caught specimens. c. Dominant male, the holotype. d. Female (RMCA 2019.002.P.0017; 102.1 mm SL) to illustrate the live colour patterns.
Haplochromis aquila sp. nov. c–d. Photographs of freshly caught specimens. c. Dominant male, the holotype. d. Female (RMCA 2018.008.P.0352; 108.7 mm SL) to illustrate the live colour patterns. Haplochromis kimondo sp. nov. c–d. Photographs of freshly caught specimens. c. Holotype, a dominant male. d. Female (RMCA 2018.008.P.0364; 128.1 mm SL) to illustrate the live colour patterns. Haplochromis falcatus sp. nov. c–d. Photographs of freshly caught specimens. c. Dominant male (RMCA 2017.006.P.0416; 119.1 mm SL). d. Female (RMCA 2016.035.P.0257; 112.8 mm SL) to illustrate the live colour patterns.
Haplochromis curvidens sp. nov. c–d. Photographs of freshly caught specimens. c. Holotype, an adult male. d. Female (RMCA 2018.008.P.0340); 90.2 mm SL) to illustrate the live colour patterns. Haplochromis quasimodo sp. nov. c–d. Photographs of freshly caught specimens. c. Dominant male (RMCA 2018.008.P(HP3072); 123.7 mm SL). d. Female (RMCA 2018.008.P(HP3064); 116.6 mm SL) to illustrate the live colour patterns. Haplochromis squamipinnis Regan, 1921. c–d. Photographs of freshly caught specimens. c. Dominant male (RMCA 2016.035.P.0250; 169.7 mm SL). d. Female (RMCA 2016.035.P(HP823); 129.6 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced. |
Haplochromis curvidens sp. nov.
Etymology: Specific name from the Latin ‘curvus’ for ‘curvature’, and ‘dentatus’ for ‘tooth’; referring to strongly recurved oral teeth.
Haplochromis pardus sp. nov.
EtymologySpecific name from the Latin ‘pardus’ for ‘leopard’; referring to nearly uniform black to yellow-pink flanks with clear black blotches, i.e., interrupted horizontal and vertical stripes.
Haplochromis quasimodo sp. nov.
Etymology: Specific name from Quasimodo, hunchbacked character in Victor Hugo’s novel ‘Notre-Dame de Paris’ (1831); referring to rather shallow head and deep and rhomboid bodies of large specimens.
Nathan Vranken, Maarten Van Steenberge, Annelies Heylen, Eva Decru and Jos Snoeks. 2022. From A Pair to A Dozen: The Piscivorous Species of Haplochromis (Cichlidae) from the Lake Edward System. European Journal of Taxonomy. 815(1), 1-94. DOI: 10.5852/ejt.2022.815.1749
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