Сообщество, посвящённое ра Below are the 2 most recent journal entries recorded in the "Сообщество, посвящённое ра" journal:
April 6th, 2014
10:05 pm
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Mahajangasuchus

 Mahajangasuchus is an extinct genus of crocodyliform which had blunt, conical teeth. The type species, M. insignis, lived during the Late Cretaceous; its fossils have been found in the Maevarano Formation in northern Madagascar. It was a fairly large predator, measuring up to 3 m (10 ft) with a weight up to 360 kg (800 lbs).

 Sereno et al.. (2001) placed the genus within the family Trematochampsidae, although a more recent study by Turner and Calvo (2005) placed it within Peirosauridae. It was placed in the newly constructed family Mahajangasuchidae along with the genus Kaprosuchus by Sereno and Larrson (2009).


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06:23 pm
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Kaprosuchus

 Kaprosuchus is an extinct genus of mahajangasuchid crocodyliform. It is known from a single nearly complete skull collected from the Upper Cretaceous Echkar Formation of Niger. The name means "boar crocodile" from the Greek kapros ("boar") and souchos ("crocodile") in reference to its unusually large caniniform teeth which resemble those of a boar. It has been nicknamed "BoarCroc" by Paul Sereno and Hans Larsson, who first described the genus in a monograph published in ZooKeys in 2009 along with other Saharan crocodyliformes such as Anatosuchus and Laganosuchus. The type species is K. saharicus.

 Kaprosuchus is known from a nearly complete skull in which the lower jaw measured 603 mm long, whilst the entire animal is estimated to have been around 6 metres (20 feet) in length. It possesses three sets of tusk-like caniniform teeth that project above and below the skull, one of which in the lower jaw fits into notches in upper jaw. This type of dentition is not seen in any other known crocodyliform. Another unique characteristic of Kaprosuchus is the presence of large, rugose horns formed from the squamosal and parietal bones that project posteriorly from the skull. Smaller projections are also seen in the closely related Mahajangasuchus.

 The snout of Kaprosuchus shows generalized proportions and the naris is positioned dorsally. In Kaprosuchus many teeth are hypertrophied and labiolingually (laterally) compressed, unlike those of crocodyliforms with similarly shallow snouts, which are usually subconical and of moderate length. Another difference between the skull of Kaprosuchus and those of crocodyliforms that also possess dorsoventrally compressed snouts is the great depth of the posterior portion of the skull.

 In Kaprosuchus, the orbits (i.e., eye sockets) open laterally and are angled slightly forward rather than upward. The orbits turned forward suggest that there was somewhat stereoscopic vision, i.e., an overlap in the visual field of the animal.

 The surfaces of the premaxillae are rugose with the edges elevated above the body of the bone, suggesting that a keratinous shield would have been supported by the rugosities at the tip of the snout. Along the interpremaxillary suture, the area where the two premaxillae meet, the surface is smooth, giving the paired rugosity of the premaxillae the resemblance of a moustache in anterior view.

 Kaprosuchus is a member of the family Mahajangasuchidae along with closely related Mahajangasuchus insignis from the Upper Cretaceous of Madagascar. Although it differs greatly from any other known crocodyliform, Kaprosuchus shares several characteristics with Mahajangasuchus. These include the obliteration of all but the posterior portion of the internasal suture; a laterally facing rugose external articular fossa; the positioning of the jaw joint below the posterior maxillary teeth; a deep, anterodorsally oriented mandibular symphysis; a vertically descending ectopterygoid that is slightly inset from the lateral margin of the jugal; a flared choanal septum forming an articular foot for the palatine; and the hornlike dorsal projection of the external rim of the squamosal (although this is much more developed in Kaprosuchus than Mahajangasuchus).

 Kaprosuchus is thought to have been a primarily if not exclusively terrestrial predator. Evidence for this behavior includes the positioning of the orbits laterally and somewhat anteriorly, which suggests an overlap in vision. This is unlike many other neosuchians, including extant crocodilians, in which the orbits are positioned dorsally as an adaptation to aquatic predation where the head can be held underwater while the eyes remain above the surface.

 Additional support for terrestrial predation can be found in the teeth and jaws. The enlarged caniniforms are sharp-edged and relatively straight, unlike the fluted, subconical, recurved teeth of aquatic crocodyliforms. Because the retroarticular process of the lower jaw is long, it is likely that the jaws were able to open relatively quickly with a large gape to allow for the opposing caniniforms to clear one another. The fused nasal bones are thought to have provided reinforcement for the jaws against compression associated with a powerful bite. The telescoped, dorsally positioned external nares are seen as protection against impact if the animal rammed prey with its robust snout. The keratinous shield thought to have covered the tip of the snout would have provided further protection.

 Mahajangasuchidae is an extinct family of neosuchian crocodyliforms. It currently contains two genera, Mahajangasuchus and Kaprosuchus, both of which lived during the Late Cretaceous in Gondwana. It is defined as the most inclusive clade containing Mahajangasuchus insignis but not Notosuchus terrestris, Simosuchus clarki, Araripesuchus gomesii, Baurusuchus pachecoi, Peirosaurus torminni, Goniopholis crassidens, Pholidosaurus schaumbergensis, or Crocodylus niloticus. Phyogenetically, Mahajangasuchidae is placed just outside pholidosaurids and more derived neosuchians.

 Defining characters of the family include fused nasals, a jaw articulation below the posterior maxillary tooth row, a deep mandibular symphysis that is oriented anterodorsally, and the formation of a hornlike posterodorsal process from the squamosal and parietal (which is much more pronounced in Kaprosuchus).


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