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https://www.science.org/doi/10.1126/science.abq5158

Fundamentals of empathy
Emotional contagion, in which individuals display fear or distress behaviors in response to observations of the same in another, is considered a basal form of empathy and is known to occur in fishes. Akinrinade et al. have shown that the neuropeptide oxytocin is responsible for these behaviors in zebrafish, as it is in mammals (see the Perspective by DeAngelis and Hofmann). They also found that the same regions of the brain are involved in zebrafish and in mammals. Such homologies in emotional response mechanisms across fishes and mammals suggest that this most basal form of empathy could have evolved many, many millions of years ago.

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